ScienceDirect - Current Biology : Hyena societies

Current Biology : Hyena societies: "Although the family Hyaenidae contains only four living species, and is therefore one of the smallest among mammalian carnivores, it is remarkable for its ecological and social diversity, which surpasses that of much larger families such as canids and felids. The four extant hyena species represent the last remnants of a family that was once large and diverse"